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Miki's feelings for his sister are terribly complicated and ambivalent. On the one hand, she repulses him with her bitterness, cynicism, sexual immodesty, and darkness. On the other, she draws and possesses him by a memory of what they were together, guilt, shared experiences, and the affection a brother has for a sister.
In many ways, the twins form opposites in the fashion of the ying-yang tradition. Miki is a dreamer and a romantic, always seeking to find his shining thing, while Kozue is a cynic and perhaps even a nihilist, believing everything but herself and her brother to be dirty. Miki is primarily shy though still very popular, but Kozue seems to have a lot of admires with whom she is very friendly (even if she secretly despises them). Miki's sexual development appears to be about that of a boy his age, while Kozue appears to have significantly more experience. Miki internalizes his feelings, but Kozue lashes out with them, even if she does so indirectly. Miki is a reserved, self-doubting genius and Kozue is a proud, self-destructive second hander.
Even so, they share a close bond by being family. Their parents appear disappointing, although Kozue is more vocal about her disdain than her brother. Perhaps they always lumped the twins together and failed to pay attention to them unless they had something to gain, as with their supposed piano talent. Perhaps they were hurt in the separation of their parents or maybe the events that lead up to the separation were painful. At any rate, they (but especially Kozue) feel a sense of abandonment, as symbolized by the birds for whom they build a house.
Also there was the time they spent together in the garden. Miki claims that is the only time he was ever happy, which suggests a rather unhappy childhood (supporting the idea of an unsympathetic family). Kozue had all of her brother's attention and love, not to mention praise and admiration that she did not deserve. Miki had his muse and Kozue had her brother. Depending on how accurate the above inferences about their family is, perhaps all they had was each other.
Of course, Kozue also holds him with guilt. He blames himself for her deciding she doesn't want to play the piano, and while he mourns for the loss of his muse, he also mourns the loss of the beautiful music she could have made. He still doesn't realize that it was all a trick. This, no doubt, influences his decision to fight for Anthy for the sake of protecting her beautiful music.
Still, Miki's feelings for her are rather ambivalent. He worries about her and her reckless behavior and he certainly seems love very deeply, otherwise her efforts to hurt him wouldn't be so effective. On the other hand they rarely seem to have anything pleasant to say to each other. When he carries her home after she falls into his arms while saving the birds, Miki complains that she is sticky and he doesn't want her to touch him too much. Whatever disdain and disappointment he feels concerning his sister, he will always love her. The problem is that he must love her and not an illusion of her.
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